New data bring to 18 the number of troglobitic Ptomaphagus species in the SE USA. The new species are Ptomaphagus chromolithus and P. torodei; full species rank is given to P. solanum, P. fecundus, P. longicornis and P. julius; P. laticornis is resurrected from synonymy. A phylogeny is proposed. A model of speciation is proposed with cave populations isolated during interglacials, and with probable overland dispersal and interpopulation contact during glacials. -from Author
Canadian Journal of Zoology
Department of Biology

Peck, S. (1984). The distribution and evolution of cavernicolous Ptomaphagus beetles in the southeastern United States ( Coleoptera: Leiodidae: Cholevinae) with new species and records. Canadian Journal of Zoology, 62(4), 730–740. doi:10.1139/z84-103